Specter is the main antagonist throughout the Ape Escape games. He is best known for his advanced Peak Point Helmet and hoverchairs that he uses to accomplish his evil schemes. He drives the Peak Point Tram, a transport that it's rail streches from New Freezeland to Mysterious Age.
Specter was an ordinary white monkey who performed at the Monkey amusement park until he found one of the Professor's inventions "The Peak Point Helmet", an invention that increases the intelligence of the brain of whoever wears it. After putting this strange item on his head it took control of his mind turning him into a power mad monster. He leads the Pipo Monkeys.
Specter appears alongside Spike and lots of Pipo Monkeys. Specter challenges the Heroes to battle his army of Pipo Monkeys. The heroes defeat all of them and Spike heals the monkeys. Specter, Spike and the Monkeys join.